Phylogenetic relationships of subspecies were examined by means of 16S rRNA gene sequencing. Sequence similarities among subsp. strains exceeded 99·0%, but values among subsp. strains ranged from 96·4 to 100%. Sequence similarites between strains representing the two different subspecies ranged from 95·7 to 99·0%. An intervening sequence was identified in certain of the subsp. strains. strains occupied two distinct branches in a phylogenetic analysis of the genus , emphasizing the need for multiple strain analysis when using 16S rRNA gene sequence comparisons for taxonomic investigations.


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