Outer membrane vesicles and fragments from , and were isolated and examined semiquantitatively for 19 enzyme activities by using the API ZYM micromethod. The enzyme contents of vesicles and fragments were compared with the enzyme contents of whole cells of the same organisms. Enzymic data were analyzed by using principal-component analysis and soft independent modeling of class analogy. This technique allowed us to distinguish among the closely related organisms , and was divided into two groups of strains. fell outside or on the border of the class. , and fell outside the , and classes.


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