Analysis of the nodulation characteristics of transposon-induced mutants of bv. revealed that and the closely-linked genes contribute to nodulation of peas () and the vetch Although mutation of alone had no significant effect on nodulation of either legume, a double mutant lacking both and nodulated both legumes very poorly. Similarly, a double mutant lacking and either or nodulated peas less efficiently than a mutant. Thus, although mutations affecting only the genes normally have no observed effect on nodulation, these genes do appear to contribute to pea nodulation. When transferred to a wild-type strain of bv. , neither nor the gene region conferred pea or vetch nodulating ability. However, in a mutant of bv. did confer a significant level of vetch nodulating ability, indicating that the secreted NodO protein can play a role in determining legume recognition by bv.


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