Gene deletions of the operon of showed that the and genes, but not , were required for formation of wild-type levels of the LHII complex. Deletion of or also impaired photosynthetic growth. The effects of deletion were suppressed by secondary mutations that mapped outside the operon. Fusion of a gene to ' showed that most of transcription originated from upstream of . RNA blot analysis revealed a 2.4 kb transcript that hybridized to probes specific for the and regions, indicating that some operon messages extend from just before the gene to just after the gene.


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