Part A of Appendix 9 – Orthography of the International Code of Nomenclature of Prokaryotes regulates the formation of compound generic names and specific epithets derived by combining two or more words or word elements of Latin and/or Greek origin, using the word stems and connecting vowels (-o- or -i-) following word elements derived from Greek and Latin, respectively. The rules given and the exceptions listed are suitable for substantives (nouns) and adjectives used as word elements, but not for prepositions and prefixes. Therefore, we propose a non-retroactive modification of Appendix 9 so that the guidelines given in Part A apply only to compound names that include a noun or an adjective in a non-final position. We also propose guidelines for the proper use of Greek and Latin prepositions, prefixes and adverbs in compound names in which the following word element starts with a vowel.


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  1. Parker CT, Tindall BJ, Garrity GM. International code of nomenclature of prokaryotes. prokaryotic code (2008 revision). Int J Syst Evol Microbiol 2019; 69:S1–S111
    [Google Scholar]
  2. Trüper HG, Euzéby JP. International code of nomenclature of prokaryotes. Appendix 9: Orthography. Int J Syst Evol Microbiol 2009; 59:2107–2113 [View Article][PubMed]
    [Google Scholar]
  3. Turland NJ, Wiersema JH, Barrie FR, Greuter W et al. (editors) International Code of Nomenclature for Algae, Fungi and Plants (Shenzhen Code) Adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017, Regnum Vegetabile 159. Glashütten; Glashütten Koeltz Botanical Books: 2018
    [Google Scholar]
  4. Goodwin WW. A Greek Grammar Boston: Ginn & Company; 1900
    [Google Scholar]
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