A total of 219 strains belonging to the genera and were studied; 171 of these strains were isolated from mollusks, mainly snails and slugs, obtained from around the world. On the basis of DNA-DNA hybridization data, the strains were grouped into 11 genomospecies. A total of 44 phenotypic characters were used to differentiate the genera and at the genus level and to identify genomospecies. There were significantly higher phenotypic probability distances between the genomospecies in the genus and the genomospecies in the genus than between the genomospecies in the same genus. Therefore, the existence of and as different genera was confirmed. The existence of new species necessitated broadening the definitions of both genera. In two cases, two species could not be quantitatively differentiated biochemically, and several other pairs of species could be separated only by the results of one biochemical test. Nonetheless, combinations of several characteristics were used to differentiate all of the species with levels of certainty ranging from log 10.79 to log 57.77 (calculated as probability distances). The following new species are proposed: (type strain, ATCC 51602 [DSM 9390]), (type strain, ATCC 51604 [DSM 9393]), (type strain, ATCC 51605 [DSM 9396]), (type strain, ATCC 51606 [DSM 9397]), (type strain, ATCC 51607 [DSM 9401]), (type strain, ATCC 51608 [DSM 9404]), (type strain, ATCC 51609 [DSM 9406]), and (type strain, ATCC 51603 [DSM 9409]).


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