The 16S rRNA gene sequences of 30 strains representing 23 validated species and 7 currently non-valid species were determined. These sequences were aligned with the sequences of other species and related actinomycete taxa. A comparative sequence analysis revealed that there is considerable phylogenetic depth and internal structure in the genus and the amycolate species were located at the periphery of the genus . It was evident that the species of the genus form a monophyletic association and, together with other chemotype IV and mycolic acid-containing taxa (including the genera , and ), form a natural suprageneric group.


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