The taxonomic relationships among nonpigmented species of the genus were assessed by DNA-DNA hybridization. The species formed two groups based on DNA homology values and moles precent G+C of DNA: , the type species, had a moles percent G+C content 4 to 7% higher than those of the other group, and it had less than 5% DNA homology to the second group formed by the remaining species. The remaining species formed seven DNA homology subgroups, of which five corresponded to the type strains of , and . A sixth subgroup included the type strains of and , which showed hybridizations of more than 80%. The species in the second group were related to each other at the genus level, but a nomenclatural proposal is postponed. The hybridization data showed that the name Simidu et al. 1990 should be recognized as a junior subjective synomyn of (ZoBell and Upham 1944) Reichelt and Baumann 1973. The seventh subgroup included two as yet unassigned strains which were misidentified as to species.


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