Using a variety of physiological, biochemical, and molecular systematic analyses, we have shown previously that there are four groups within the species . Two of these groups of strains correspond to the recently proposed taxa subsp. and subsp. . In this paper we show that the two remaining groups are distinct and formally propose that they should be recognized as subsp. (type strain, NCTC 11326) and subsp. (type strain, NCTC 12276). The tests which we used did not allow a full assessment of the status of subsp. compared with subsp. .


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