The phylogenetic interrelationships of members of the genus were investigated by using reverse transcriptase sequencing of 16S rRNA. The sequence data indicate that at the intrageneric level the genus consists of the following two closely related but distinct lines of descent: (i) the group of species (including , and ) and (ii) the species and . At the intergeneric level a specific phylogenetic relationship between the genera and was evident. The sequence data clearly demonstrated that the genus is phylogenetically remote from the genus and should not be included in an extended family .


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