A total of 15 strains of the genus were characterized chemotaxonomically and physiologically in an attempt to resolve their taxonomy. These strains fell into two taxa, which differed in cell wall composition and menaquinone, phospholipid, and fatty acid patterns. Ten of the strains assessed in this study belong to the genus and can be assigned to five species on the basis of their physiology. subsp. is heterogeneous; remains centered on the type strain (strain DSM 43111), and the remaining strains form a new subspecies, subsp. sp. nov. (type strain, strain DSM 43377). The former subsp. is recognized as a close relative of the latter organism and is designated subsp. comb. nov. Two formerly misclassified species of the genus not on the Approved Lists are revived as sp. nov. and sp. nov. The remaining five species (, and ) appear to be misclassified, and, on the basis of their similarities to the type species , reclassification in the genus should be considered. and were not included in this study.


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