Deoxyribonucleic acid hybridization (hydroxyapatite method, 55 and 70°C) was used to characterize 38 serovars from 22 named serogroups of and , 6 serovars from 4 new unnamed serogroups of , and single serovars of the proposed species and Deoxyribonucleic acid relatedness confirmed the validity of and The well-accepted species and , as currently defined, were extremely heterogeneous. Relatedness results revealed at least five new species among the parasitic serovars formerly included in and two new species among the saprophytic serovars formerly included in Serogrouping did not equate with species identification, as serovars from several different subserogroups belonged to different species. The new species named in this paper are , and .


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