The varietal relationship within the species was determined by a study of deoxyribonucleic acid relatedness and by a comparison of the electrophoretic mobilities of 12 metabolic enzymes. The nuclear genomes of var. and var. showed about 28% relative binding to hydroxylapatite in intervarietal deoxyribonucleic acid reassociation experiments, and the allozymes encoded by eight loci were different in the two taxa. Based on these molecular criteria, as well as allopatry, different host plants, and weak sexual interaction, var. is raised to the rank of species as (Starmer et al.) Holzschu et al. comb. nov.


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