We describe and , two species that have as their habitats necrotic tissues of spp. and spp., respectively. , 21 strains of which were isolated, is homothallic and occurs in nature exclusively in the diploid state. It produces asci with two hat-shaped spores, which are rapidly released upon maturity. This species is nonfermentative and assimilates few carbon compounds. The guanine-plus-cytosine content range of the nuclear deoxyribonucleic acid (eight strains) is 27.4 to 28.4 mol%, and the average ± standard deviation for eight strains is 27.8 ± 0.4 mol%. , 48 strains of which were isolated, has the same phenotypic properties and deoxyribonucleic acid base composition as , but lacks the ability to produce ascospores and is resistant to triterpene glycosides in growth media. The deoxyribonucleic acids of and show more than 96% homology, but the two species are separated geographically and by host plant. occurs in species in southern Arizona and Texas, whereas is found almost exclusively in columnar cacti of the genus on certain Caribbean islands and in Baja California, Mexico. The type strain of is strain UCD-FST 83-467.3 (= ATCC 58091 = CBS 7119), and the type strain of is strain UCD-FST 76-355A (= ATCC 58088 = CBS 7121).


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