We studied 170 strains of , and related taxa. The overall similarities of the test strains were determined by comparing 120 unit characters, using the simple matching and pattern coefficients. Clustering was achieved by using the unweighted pair group method with averages. Cluster composition was not markedly affected by the coefficient used or by test error (estimated at 4.9%). The numerical data, together with results from previous chemical and genetic studies, showed that the genus is markedly heterogeneous. Strains received as fell into two large aggregate groups. One, sensu stricto, contained , and . was not studied, but other genetic data suggest that it should also be included in this group. The other aggregate group encompassed , and . The representatives of , and studied formed distinct clusters separate from both of the aggregate groups. strains formed a cluster clearly distinguished from the clusters containing the strains.


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