Proposed as neotype strains of their respective species are the following: ATCC 486, ATCC 487, ATCC 484, ATCC 482, ATCC 488, ATCC 489, and ATCC 491. Deoxyribonucleic acid-deoxyribonucleic acid reassociation studies on these and three other reference strains of clearly revealed seven distinct species: , and ATCC 489 is identical with ATCC 488 on the basis of genetic evidence. These results are confirmed by physiological data. is therefore a junior synonym of . The polysaccharide and amino acid composition of the cell wall, the electrophoretic mobility of the L-lactate dehydrogenase, and the utilization of certain sugars and organic acids were found to be useful characters in species differentiation. Although the deoxyribonucleic acid-deoxyribonucleic acid homology values within the genus range between 20 and 100%, those between strains of and strains of , and are lower than 8%. These data, supported by the high guanine plus cytosine content of the DNA, the ability to decompose cellulose, and the similarity in the peptidoglycan types, demonstrate the coherence of the members of the genus and the validity of the inclusion of this genus in the family .


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