Eleven strains of , four strains of pathovar , six strains of subsp. , two strains of , and one sp. were compared by physiological, biochemical, serological, and pathological tests. All strains tested utilized sucrose, glucose, fructose, glycerol, and caprate as sole carbon sources, hydrolyzed Tween 80, reduced litmus milk, and grew at 0°C. All strains were arginine dihydrolase and oxidase negative and all strains failed to hydrolyze starch, reduce nitrate, or grow at 41°C. Cells of all strains were immunofluorescent positive when stained by the indirect immunofluorescence test with antiserum to cells of or subsp. . No differences among strains were detected in test tube agglutination tests. No significant differences were detected in the susceptibility of (oats), (rye), (wheat), (barley), (smooth bromegrass), (Japanese brome), (chess brome), (cheatgrass), (quackgrass), (maize), or (timothy) to the different strains. All strains produced toxin in Wooley medium except and strains of subsp. from Italian ryegrass. The present division of strains of based on minor differences in pathogenicity and symptomology is untenable.


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