A new species of the genus has been recovered 38 times in the Sonoran Desert from “rot pockets” of cereoid cacti and from species which utilize the cacti. We have named the species due to its absolute requirement for methionine or cysteine. is heterothallic and demonstrates an agglutination reaction when opposite mating types are mixed. Two varieties are designated based on the combination of mannitol assimilation and ecological habitat. var. , the type variety, was recovered from cacti in the subtribe and cannot assimilate mannitol, whereas var. was found in cacti of the subtribe and can assimilate mannitol. Results are given which demonstrate that the assimilation of mannitol is controlled by a single genetic locus. An evaluation of the interfertility and postmating viability among the two varieties and possibly identical organisms was made. The base composition of the nuclear deoxyribonucleic acid (average of 10 strains) is 33.05 ± 0.19 mol% guanine plus cytosine. The type strain of and of the type variety, var. , is UCD-FST 76-401B (=ATCC 36080 = CBS 6940). The type strain of var. is UCD-FST 76-384A (=ATCC 36079 = CBS 6943).


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