Relatedness among strains of was determined by using interspecific deoxyribonucleic acid reassociation followed by chromatography on hydroxyapatite. Relatedness was also assessed among and representatives from all other established genera of the family Twenty strains of from five countries and 11 states, isolated from diseased human, normal human, and invertebrate sources, all formed one highly related hybridization group. exhibited 20 to 29% relatedness to other enterobacteria. These data are consistent with the previous designation of a separate tribe, , for a single genus () and species ().


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