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Budding as a mode of replication of nitrite oxidizing bacteria was observed and illustrated by Winogradsky in his work on Nitrobacter, and, shortly thereafter, by Stutzer and Hartleb, studying Nitromicrobium germinans. The identity of this bacterium with the Nitrobacter cultures of Winogradsky was established by Stutzer. Nitrorcrobium is thus a later synonym to Nitrobacter. The species name germinans would have had priority over winogradsky but it was overlooked. Opinions 20 and 23 of the Judicial Commission clearly validate Nitrobacter winogradskyi Winslow et at. 1917, and thus Nitromicrobium germinans is to be placed among the synonyms of N. winogradskyi.
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